>Port forwarding on the client router is set to 443 on 192.168.2.5.
Forgive me for a simplistic suggestion.
Are you sure your router is set up so that it receives requests at port 4431 and forwards to the port 443 of your destination IP?
I just found this:
https://www.canyouseeme.org/
NOT suggesting you use this, so please use it at your own risk!